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RE: Haha, even more questions



Tom Holtz wrote:
 
> B) Almost all recent studies ally troodontids with dromaeosaurids and birds
> rather than with ornithomimosaurs, so the arctometatarsus is clearly
> convergent between these groups.
 
The arctometatarsus may have evolved at many as *six* times in theropod evolution: ornithomimids, tyrannosaurids, mononykines, elmisaurids, avimimids, and troodontids.  However, it is probable that two or more of these groups share a common ancestry for the arctomet foot - elmisaurids (?=caenagnathids)+avimimids, and ornithomimosaurs+tyrannosaurs, for example.  Also, the arctomet foot (or an incipient version of it) might have arisen deep in coelurosaur phylogeny and was subsequently refined or lost in individual theropod lineages.  Possibilities abound.
 
 
> > 4) Is Confuciusornis dui still considered a valid species of Confuciusornis?
>  I'll get out my species-ometer and check.  --whoop whoop 
> whoop--, Ah, well, inconclusive... :-)
Chiappe et al.'s (1999) species-meter came to the same conclusion (or same lack of conclusion).  They do not affirm or deny the validity of _C. dui_ (i.e. whether or not it is a synonym of _C. sanctus_).  The most distinctive feature of _C. dui_ is (said to be) the upturned Woody Woodpecker-like beak evident in the keratinous outline (not the bone underneath, though).  Reading between the lines: (a) the shape of the beak (cornified rhamphotheca) may not be unique to the _C. dui_ type specimen (IVPP V11553) and may be present in _C. sanctus_ too; (b) the unusual postcranial features ascribed to IVPP V11553 may prove to be absorbed by the broad intraspecific variation reported for _C. sanctus_.
 
 
References
 
Chiappe, L.M., Ji S.-A.; Ji Q., and Norell, M.A. (1999). Anatomy and systematics of the Confuciusornithidae (Theropoda: Aves) from the Late Mesozoic of Northeastern China.  Bulletin of the American Museum of Natural History 242: 89pp.  (A good read)
 
 
 
Tim
 
 
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Timothy J. Williams, Ph.D.

USDA-ARS Researcher
Agronomy Hall
Iowa State University
Ames IA 50014

Phone: 515 294 9233
Fax:   515 294 9359