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Re: Apomorphy-based definitions (of mammals [...])
Yes, I was being imprecise last night, and David was right to point
this out (shame on me). Total loss of the last vestiges of what was once
the reptilian jaw articulation occurred at the Hadrocodium stage. However,
the mammalian jaw joint had long since taken over as the functional jaw
joint.
That the latter had occurred in Adelobasileus can only be inferred at
the present time, so the definition was made more precise with its
remarkable "incipient promontorium". Once we have more complete fossil
material of Adelobasileus and/or its closest relatives, a more precise
definition will be possible without having to invoke controversial
inferences (forced on us by incomplete material).
So getting back to Aves, this is why I am so concerned that the
definition of the "true" semilunate be made more precise by including
evolutionary transitions in one or more of the adjacent bones. This is what
I mean by learning from the mammalian equivalent, and try to do it better
with Aves. Such a precise definition would presumably even convince the
most staunch Feducciaries that birds clearly evolved from coelurosaurian
dinosaurs (but bringing that debate into sharper focus would be just icing
on the cake, not the major reason for doing it). I have given this a great
deal of thought, and I'm convinced it is the best way to proceed (regardless
of whether you want to call Aves a major "Class" or a major "Clade"---to a
cladisto-eclecticist, Aves is both a Clade *and* a Class).
------ Ken
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