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Re: Apomorphy-based definitions (of mammals [...])




Yes, I was being imprecise last night, and David was right to point this out (shame on me). Total loss of the last vestiges of what was once the reptilian jaw articulation occurred at the Hadrocodium stage. However, the mammalian jaw joint had long since taken over as the functional jaw joint.
That the latter had occurred in Adelobasileus can only be inferred at the present time, so the definition was made more precise with its remarkable "incipient promontorium". Once we have more complete fossil material of Adelobasileus and/or its closest relatives, a more precise definition will be possible without having to invoke controversial inferences (forced on us by incomplete material).
So getting back to Aves, this is why I am so concerned that the definition of the "true" semilunate be made more precise by including evolutionary transitions in one or more of the adjacent bones. This is what I mean by learning from the mammalian equivalent, and try to do it better with Aves. Such a precise definition would presumably even convince the most staunch Feducciaries that birds clearly evolved from coelurosaurian dinosaurs (but bringing that debate into sharper focus would be just icing on the cake, not the major reason for doing it). I have given this a great deal of thought, and I'm convinced it is the best way to proceed (regardless of whether you want to call Aves a major "Class" or a major "Clade"---to a cladisto-eclecticist, Aves is both a Clade *and* a Class).
------ Ken




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