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Mass distribution and running



Hello.
 
Quadrupedal mammals have most of their mass supported by their forelimbs. Thanks to this, their center of mass is fell forward of the hips, and when the animal runs, the force resulting from the interaction between the hindlimbs and ground that goes through the center of mass has a relatively long horizontal component, which is good from the point of running.
 
However, dinosaurs have most of their mass supported by their hindlegs. It is pretty obvious that this has to be so in the bipedal forms, but this is also true of the quadrupedal ones. I wonder why. Wouldn't it have been better for the four-limbed herbivores to have their center of mass farther forward?
 
It has been proposed that the peculiar distribution of mass in herbivorous dinosaurs was due to their descent from bipedal forms. However, it doesn't seem very plausible to me that this would have kept them from evolving into more efficient forms. So, probably they were already just as efficient as quadrupedal mammals. But how could this be?
 
Henri Rönkkö