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Re: Pterosaur Help
In a message dated 12/4/01 7:53:23 PM EST, mhanson54@attbi.com writes:
<< All that is now left is Nyctosaurus, Cycnorhamphus, Dimorphodon, Tupuxuara
and Domeykodactylus, and Doratorhynchus. >>
Parafamily: DIMORPHODONTIDAE (nom. emend. ex Dimorphodontae) Seeley, 1870
Census: 2 genera, 3 species
= Dimorphodontinae Hooley, 1913
= Dimorphodontidae Nopcsa, 1923
Genus: Dimorphodon Owen, 1859
= Dimorphodon Owen, 1858 [nomen nudum]
D. macronyx (Buckland, 1829) Owen, 1859â
= Pterodactylus macronyx Buckland, 1829
= Dimorphodon macronyx (Buckland, 1829) Owen, 1858 [nomen nudum]â
= Rhamphorhynchus macronyx (Buckland, 1829) von Meyer, 1857 (as Pterodactylus
(Rhamphorhynchus) macronyx)
= Pterodactylus (Dimorphodon) macroynx Owen, 1874 [sic]
= Pterodactylus mardeni Owen, 1874 [sic]
= Pterodactylus marderi Owen, 1874 [nomen dubium]
D. weintraubi Clark, Hopson, HernÃndez, Fastovsky & Montellano, 1998
Genus: Peteinosaurus Wild, 1978
= Peteinosausus Chaline, 1990 [sic]
P. zambellii Wild, 1978â
Genus: [To be described from the Dockum Formation of Texas; S. Chatterjee,
pers. comm.]
Family: GALLODACTYLIDAE Fabre, 1974
Census: 2 genera, 3 species
= Gallodactylinae Fabre, 1981
Genus: Cycnorhamphus Seeley, 1870
= Cycnorhamphys Fabre, 1981 [sic]
= Cygnorhamphus Seeley, 1870 [sic]
= Cynorhamphus Romer, 1966 [sic]
= Gallodactylus Fabre, 1974
C. suevicus (Quenstedt, 1855) Seeley, 1870â
= Pterodactylus suevicus Quenstedt, 1855 non Oken, 1825
= Gallodactylus suevicus (Quenstedt, 1855) Fabre, 1974
= Pterodactylus wuerttembergicus Quenstedt, 1854 [nomen dubium]
= Pterodactylus wÃrttembergicus Quenstedt, 1854 [nomen dubium]*
= Pterodactylus eurychirus Wagner, 1857
= Pterodactylus (Ornithocephalus) eurychirus (Wagner, 1857) Wagner, 1857
= Ornithocephalus eurychirus (Wagner, 1857) Olshevsky, 1978
= Pterodactylus suevicus eurychirus Wagner, 1858
= Pterodactylus eurycheirus Newton, 1888 [sic]
= Pterodactylus wurtembergicus Fabre, 1981 [sic]
C. canjuersensis (Fabre, 1974) Bennett, 1996
= Gallodactylus canjuersensis Fabre, 1974â
NOTE: The species in this genus are organized according to Fabre (1974) and
Bennett (1996).
Genus: Diopecephalus Seeley, 1871
= Diocephalus Fabre, 1981 [sic]
D. longicollum (von Meyer, 1854) Seeley, 1871â
= Pterodactylus longicollum von Meyer, 1854
= Gallodactylus longicollum (von Meyer, 1854) Fabre, 1974
= Pterodactylus longipes MÃnster, 1836 [nomen oblitum]
= Ornithocephalus longipes (MÃnster, 1836) Olshevsky, 1978 [nomen oblitum]
= Pterodactylus (Ornithocephalus) vulturinus Wagner, 1857
= Pterodactylus vulturinus (Wagner, 1857) Wagner, 1858
= Ornithocephalus vulturinus (Wagner, 1857) Olshevsky, 1978
= Pterodactylus longicollis Wagner, 1858 [sic]
= Pterodactylus suevicus O. Fraas, 1878 non Oken, 1825 non Quenstedt, 1855
= Cycnorhamphus fraasii Seeley, 1891
= Cycnorhamphus fraasi (Seeley, 1901) Seeley, 1901
= Pterodactylus fraasi (Seeley, 1901)
NOTE: The species Cycnorhamphus fraasii was originally described, briefly, in
Seeley's 1891 paper on pterosaur pelves. Seeley refigured it (with the
trivial name respelled) in his 1901 monograph Dragons of the Air. This
document is often listed, incorrectly, as the original source of the species
(e.g., in Wellnhofer, 1970 and Fabre, 1981).
Family: NYCTOSAURIDAE Nicholson & Lydekker, 1889
Census: 1 genus, 4 species
= Nyctodactylinae Williston, 1897
= Nyctosaurinae F. Plieninger, 1907
Genus: Nyctosaurus Marsh, 1876
= Nyctisaurus Marsh, 1880 [sic]
= Nyctodactylus Marsh, 1881
= Nytosaurus Miller, 1972 [sic]
N. gracilis (Marsh, 1876) Marsh, 1876â
= Pteranodon gracilis Marsh, 1876
= Nyctodactylus gracilis (Marsh, 1876) Marsh, 1881â
= Pteranodon (Nyctosaurus) gracilis (Marsh, 1876) Miller, 1972â
= Pteranodon comptus Marsh, 1876 [nomen dubium; in part]
= Nyctosaurus leptodactylus Williston, 1903
= Pteranodon (Nytosaurus) gracilis Miller, 1972 [sic]
N. nanus (Marsh, 1881) Schoch, 1984
= Pteranodon nanus Marsh, 1881
N. lamegoi Price, 1953
= Nyctodactylus lamegoi (Price, 1953) Kuhn, 1961
= Pteranodon (Nyctosaurus) lamegoi (Price, 1953) Miller, 1972
N. bonneri (Miller, 1972)
= Pteranodon (Nyctosaurus) bonneri Miller, 1972
= Pteranodon bonneri (Miller, 1972) Miller, 1972
= Nyctosaurus sternbergi Bonner, 1964 vide Miller, 1972 [nomen manuscriptum]
NOTE: See also note for Pteranodon. Nyctosaurus bonneri is probably a junior
subjective synonym of Nyctosaurus gracilis (Bennett, 1991, 1994).
Family: DSUNGARIPTERIDAE Young, 1964
Census: 5 genera, 6 species (1 doubtful)
= Dsungaripteroidea Young, 1964
Genus: Domeykodactylus Martill, Frey, Chong Diaz & Bell, 2000
D. ceciliae Martill, Frey, Chong Diaz & Bell, 2000â
Genus: Dsungaripterus Young, 1964
= Dzungaripterus Young, 1971 [sic]
D. weii Young, 1964â
= Dzungaripterus weii Young, 1971 [sic]
?D. brancai (Reck, 1931) Bakhurina, 1982 [nomen dubium]
= Pterodactylus brancai Reck, 1931 [nomen dubium]
Genus: Noripterus Young, 1973
N. complicidens Young, 1973â
Genus: Phobetor Bakhurina, 1986/KrÃyer, 1844
= Fabeter Ivakhnenko & Korabelnikov, 1987 [sic]
P. parvus (Bakhurina, 1982) Bakhurina, 1986â
= Dsungaripterus parvus Bakhurina, 1982
= Fabeter parvus Ivakhnenko & Korabelnikov, 1987 [sic]
NOTE: The above genus was named without formal description in a review
article (Priroda 1986(7): 27â 36), but it is treated as valid by Wellnhofer
(1991). It is, however, preoccupied (B. Creisler, pers. comm.) and a
replacement name will soon be published. A recent anonymous museum pamphlet
on Mongolian fossils (in Russian) illustrated a Phobetor skull and referred
it to Ornithocheiridae.
Genus: Puntanipterus Bonaparte & Sanchez, 1974
= Punctanipterus Gasparini, Leanza & Garate Zubillaga, 1987 [sic]
= Puntaniptero [Anonymous] 1976 [sic]
= Putanipterus Carroll, 1987 [sic]
P. globosus Bonaparte & Sanchez, 1974â
= Punctanipterus globosus Gasparini, Leanza & Garate Zubillaga, 1987 [sic]
Family: AZHDARCHIDAE Padian, 1986
Census: 9 genera (1 doubtful), 10 species (1 doubtful)
= Azadarchidae Varricchio, 1995 [sic]
= Azhdarchinae Nessov, January 1984
= Titanopterygiidae Padian, December 1984
Genus: Arambourgiania Nessov & Yarkov, 1989
= Arambourgiana Padian & Smith, 1992 [sic]
= Titanopterxy Frey & Martill, 1996 [sic]
= Titanopteryx Arambourg, 1959/Enderlein, 1934
A. philadelphiae (Arambourg, 1959) Nessov & Yarkov, 1989â
= Titanopteryx philadelphiae Arambourg, 1959â
= Titanopterxy philadelphiae Frey & Martill, 1996 [sic]
[New species to be described; Baird, 1986]
Genus: Azhdarcho Nessov, 1984
= Azhdarcho Nessov, 1982 [nomen nudum]
A. lancicollis Nessov, 1984â
= Azhdarcho imparidens Nessov, 1982 [nomen nudum]
Genus: Bennettazhia Nessov, 1991
B. oregonensis (Gilmore, 1928) Nessov, 1991â
= Pteranodon oregonensis Gilmore, 1928
NOTE: Murry, Winkler & Jacobs (1991) and Nessov (1991) refer Bennettazhia (or
Pteranodon) oregonensis to Azhdarchidae; S. C. Bennett (pers. comm. and 1994)
considers it a dsungaripterid.
Genus: Bogolubovia Nessov & Yarkov, 1989
= Bogoliubovia Olshevsky, 1991 [sic]
B. orientalis (Bogolubov, 1914) Nessov & Yarkov, 1989â
= Ornithostoma orientale Bogolubov, 1914 emend. Olshevsky, 1991
= Ornithostoma (Pteranodon) orientalis Bogolubov, 1914*
= Pteranodon orientalis (Bogolubov, 1914)
NOTE: Genera ending in -stoma are neuter, not feminine, and so take neuter
adjectival specific names. Hence the spelling change to Ornithostoma
orientale. Referred to the Azhdarchidae by Nessov (1991).
Genus: Doratorhynchus Seeley, 1875 [nomen dubium]
D. validus (Owen, 1870) Seeley, 1875â
= Pterodactylus validus Owen, 1870 [nomen dubium]
= Cycnorhamphus validus (Owen, 1870) [nomen dubium]
= Ornithocheirus validus (Owen, 1870) Newton, 1888 [nomen dubium]
= Ornithochirus validus Lydekker, 1888 [sic]
= Doratorhynchus validum Woods, 1891 [sic]
NOTE: This genus is not considered an azhdarchid by Nessov (1991). It is
placed in Pterodactyloidia incertae sedis by Howse & A. R. Milner (1995), who
also referred some of the material assigned to the type species to the
species Gnathosaurus macrurus.
Genus: Montanazhdarcho Padian, Horner & de Ricqles, 1994
M. minor Padian, Horner & de Ricqles, 1994â
Genus: Quetzalcoatlus Lawson, 1975
= Quetzalcoatus Nessov, 1991 [sic]
= Quetzalkoatlus Nessov & Yarkov, 1989 [sic]
Q. northropi Lawson, 1975â
NOTE: Despite numerous popular accounts of this "largest known flying
animal," this genus remains formally undescribed and is technically a nomen
nudum, but a diagnosis has been published by Nessov (1991). More than one
species may be represented in referred material from Texas.
Genus: Tupuxuara Kellner & Campos, 1988
= Tepuxuara Nessov, 1991 [sic]
T. longicristatus Kellner & Campos, 1988â
= Tepuxuara longicristatus Nessov, 1991 [sic]
T. leonardii Kellner & Campos, 1994
NOTE: Although this genus had been classified in Tapejaridae by Kellner
(1990), he now (pers. comm. at 1989 SVP annual meeting) considers it to be an
azhdarchid, based on comparison with undescribed Quetzalcoatlus skull
material at the Balcones Research Laboratory in Austin, Texas. It is referred
to Azhdarchidae by Nessov (1991).
Genus: Zhejiangopterus Cai & Wei, 1994
Z. linhaiensis Cai & Wei, 1994â
Genus: [To be described from the Maastrichtian of LaÃo, Spain; Astibia,
Buffetaut, Buscalioni, Cappetta, Corral, Estes, Garcia-Garmilla, Jaeger,
Jimenez-Fuentes, Le Loeuff, Mazin, Orue-Etxebarria, Pereda-Suberiola, J. E.
Powell, Rage, Rodriguez-Lazaro, Sanz & Tong, 1990]