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https://www.cell.com/action/showPdf?pii=S0960-9822%2820%2931511-6Feathers are the most complex integumentary structures in the animal world. They come in a variety of forms, the most familiar of which are remiges (flight feathers). Flight feathers are composed of a central shaft made up of a hollow calamus (quill), which is inserted into the skin, and a more distal rachis. Hundreds of parallel barbs branch from the sides of the rachis. In turn, smaller hooked barbules branch off the barbs, allowing them to interlock in a tight zipper-like fashion to form vanes. Variations in rachis, barb and barbule morphology result in other feather types such as contour feathers, bristles and down feathers. Feathers have a remarkable array of functions -- they form airfoils and elaborate display structures, they serve to camouflage and insulate, to generate and help detect sound, and even to disintegrate into powder to condition other feathers.