Ben Creisler
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A new ichnosite in southwest Lesotho (Upper Moyeni; Quthing District) is located within the uppermost part of the highly fossiliferous Elliot Formation, ~35 m below the conformably overlying Clarens Formation and ~65 m above the world-renowned Lower Moyeni ichnosite. While the Lower Moyeni site preserves diverse Early Jurassic ichnofossils, the ichnites at the Upper Moyeni comprise one vertebrate burrow and ~50 tridactyl tracks with footprint lengths between 15 and 51 cm. Many of the tracks preserve digital pad impressions, claw marks and displacement rims, all related to substrate conditions. The morphometric parameters of the Upper Moyeni tracks are consistent with Grallator, Eubrontes and Kayentapus. Several larger tracks with footprint lengths > 40 cm are Kayentapus-like and Eubrontes-like, and are comparable to previously described very large theropods tracks with lengths > 50 cm from the uppermost Elliot and Clarens formations. On the basis of sedimentological and ichnological evidence, the Upper Moyeni ichnofossils were formed in a palaeolandscape with small rivers and shallow lakes by burrowing tetrapods and a variety of bipedal dinosaurs (theropods), some of which were up to 7â8 m in body length. The Upper Moyeni tracks, together with the other very large tracks from coeval locations in southern Africa, collectively highlight the tendency towards increasing diversity in size of tridactyl tracks and by extension theropod trackmaker body size body size, which runs in tandem with the increasing diversity of non-sauropod, sauropodomorph body fossils in the Sinemurian-Pliensbachian of southern Gondwana.
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A new fossiliferous site from the Pinheiros-Chiniquà Sequence (Santa Maria Supersequence) with typical tetrapod components of the Dinodontosaurus Assemblage Zone (AZ) is here described. The new site is included in the Faxinal do Soturno Structural Block. The fossil content is characterized by cranial and postcranial material of rauisuchian archosaurs, traversodontid and chiniquodontid cynodonts and dicynodonts, the first occurrence of a large-sized coelacanth fish in the Brazilian Triassic, as well as ichnological material, represented by coprolites. The presence of Luangwa together with Massetognathus raises interesting questions regarding the age of this site included in the Dinodontosaurus AZ when compared with other localities of Southern Brazil, western Argentina and Africa. As in other localities of this AZ, the fossil specimens recovered in Bortolin site exhibit remarkable differences on their fossil preservation: those from the lower portion of the outcrop (distal floodplain) possess a heterogeneous preservation (variable between poor to fairly good) whereas those from the top levels (proximal floodplain) are comparatively better preserved. This new locality, with a large amount of stratigraphically controlled fossils, constitutes a key-locality to study the distribution of Triassic terrestrial tetrapods and to perform comparisons with other sites of this AZ, in which several historical records lack a precise provenance.
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