Theriosuchus morrisonensis sp. nov.
John R. Foster (2018)
A new atoposaurid crocodylomorph from the Morrison Formation (Upper Jurassic) of Wyoming, USA.
Geology of the Intermountain West 5: 287â295.
Free pdf:
A left mandible of a small crocodyliform found in the Upper Jurassic Morrsion Formation of northeastern Wyoming represents the first occurrence of the atoposaurid Theriosuchus in North America. The specimen demonstrates lower jaw morphology, including heterodonty (as indicated by alveolus shape), similar to Theriosuchus and Knoetschkesuchus, but autapomorphies and a unique combination of characters among these taxa indicate that it is a distinct, new species of Theriosuchus.
Tomasz Szczygielski, Justyna SÅowiak & Dawid DrÃÅdÅ (2018)
Shell variability in the stem turtles Proterochersis spp.Â
PeerJ 6:e6134
Free pdf:
Background
Turtle shells tend to exhibit frequent and substantial variability, both in bone and scute layout. Aside from secondary changes, caused by diseases, parasites, and trauma, this variability appears to be inherent and result from stochastic or externally induced flaws of developmental programs. It is, thus, expected to be present in fossil turtle species at least as prominently, as in modern populations. Descriptions of variability and ontogeny are, however, rare for fossil turtles, mainly due to rarity, incompleteness, damage, and post-mortem deformation of their remains. This paper is an attempt at description and interpretation of external shell variability in representatives of the oldest true turtles, Proterochersis robusta and Proterochersis porebensis (Proterochersidae, the sister group to all other known testudinatans) from the Late Triassic (Norian) of Germany and Poland.
Methods
All the available shell remains of Proterochersis robusta (13 specimens) and Proterochersis porebensis (275 specimens) were studied morphologically in order to identify any ontogenetic changes, intraspecific variability, sexual dimorphism, and shell abnormalities. To test the inferred sexual dimorphism, shape analyses were performed for two regions (gular and anal) of the plastron.
Results
Proterochersis spp. exhibits large shell variability, and at least some of the observed changes seem to be correlated with ontogeny (growth of gulars, extragulars, caudals, and marginals, disappearance of middorsal keel on the carapace). Several specimens show abnormal layout of scute sulci, several others unusual morphologies of vertebral scute areas, one has an additional pair of plastral scutes, and one extraordinarily pronounced, likely pathological, growth rings on the carapace. Both species are represented in a wide spectrum of sizes, from hatchlings to old, mature individuals. The largest fragmentary specimens of Proterochersis porebensis allow estimation of its maximal carapace length at approximately 80 cm, while Proterochersis robusta appears to have reached lower maximal sizes.
Discussion
This is the second contribution describing variability among numerous specimens of Triassic turtles, and the first to show evidence of unambiguous shell abnormalities. Presented data supplement the sparse knowledge of shell scute development in the earliest turtles and suggest that at least some aspects of the developmental programs governing scute development were already similar in the Late Triassic to these of modern forms.
pay-walled and not yet mentioned...
Wutuchelys eocenica n. gen.&Â n. sp.ÂÂ
Haiyan Tong, Julien Claude, Cheng-Sen Li, Jian Yang and Thierry Smith (2018)Â
Wutuchelys eocenica n. gen. n. sp., an Eocene stem testudinoid turtle from Wutu, Shandong Province, China.
Geological Magazine (advance online publication)
We describe here a new turtle from the early Eocene of Wutu, Shandong Province, China. This turtle with a full row of well-developed inframarginal scutes is assigned to the basalmost testudinoids while stem testudinoids were believed to disappear by the Palaeocene-Eocene boundary. This account shows that stem testudinoids crossed this boundary in their original range. The first comprehensive phylogenetic analysis of stem and modern testudinoids performed here demonstrates that the stem testudinoids, previously placed in the family âLindholmemydidaeâ, do not form a monophyletic group, and the two major clades of testudinoids (Emydidae and Geoemydidae+Testudinidae) split one from another well before the Palaeocene/Eocene boundary, prior to the Late Cretaceous.
=====