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Re: smallest ANCIENT non-bird dinosaur - was what I was asking



Had to weigh in here...

Tim Williams wrote:

Secondly, I'm not at all sure what maximization of either "safe
inferences" or
"generalizations" regarding hypothetical ancestors have to do with
clade
definitions. Does this really matter?

Given the history of paleontology and the misinformation given to the public I'd say it cannot be emphasized enough how important it is to properly gauge(and communicate) our confidence in an inference made from extant relatives. That was the logical underpinning of Witmer's EPB framework for soft tissue reconstruction, and it is equally important when framing behavioral or physiological hypotheses. For example, consider this Feduccia quote:


"Paleontologists have tried to turn Archaeopteryx into an earth-bound, feathered dinosaur. But itâs not. It is a bird, a perching bird. And no amount of âpaleobabbleâ is going to change that."

It's revealed as the steaming pile of anti-scientific tripe the moment you stop trying to extend the platonic idea of "bird" too far down the stem. The only content-filled part of that statement is that it's a perching animal (which turns out not to be correct), the rest of it is a rhetorical crutch based on an overt platonic ideal of what a "Bird" must be. Combatting this type of misinformation makes restricitng Aves to the crown group not only scientifically useful, but a valuble part of a general science education.

Don't forget that the crown-group Aves
definition not only casts _Archaeopteryx_ out of Aves, but also
_Ichthyornis_
and _Hesperornis_ as well. We may not be too confident about our
inferences
regarding_Archaeopteryx_ (including even whether it flew or not), but
we are on
much safer ground when it comes to _Ichthyornis_ and _Hesperornis_ -
both of
which would be non-avian under Gauthier and de Queiroz (2001)'s
definition.

This is the beauty of restricting names like these to crown groups; Archaeopteryx is further from the crown clade, so it's obviously more tenuous to push inferences that much further. It IS more likely that behaviors or soft tissues seen in modern birds would be found in Ichthyornis and Hesperornis, and it's intuitively obvious because they are in fact so close to birds, hence (by definition) they share more derived features. Of course you still can't assume features of (extant) birds to be plesiomorphic in Ichthyornis since they lay outside the crown group (again rendered intuitive by being close to but outside Aves), but the burden of evidence is eased relative to an animal like Archaeopteryx.

I especially like Jeff Hecht's reasoning for why this should be
avoided.)

I have cut and pasted Jeff's reasoning for ease of reference:

A general observation in reaction to this discussion -- having watched
the
public reaction to astronomers trying to redefine Pluto as a 'dwarf
planet,' I
would be wary of the public reaction to attempts to redefine birds in
a way that
leaves out Archaeopteryx. Astronomers had a fair justification for
reclassifying
Pluto, but they failed completely in justifying it to the public, who
have long
been taught Pluto was a planet, and concluded it was hair-splitting
silliness.
To the public who pay attention to such things, Archaeopteryx is the
first bird.
They would accept a flyer earlier than Archaeopteryx as a bird, but
they would
not accept trying to exclude Archaeopteryx on some arcane anatomical
or
cladistic grounds. Reclassifying something so fundamental -- and on
some level
so obvious -- could make evolution seem a less-certain science than it
is.

Personally I don't agree with this analogy. For starters, there hadn't been serious discussion of Pluto not being a planet (in the publics' eye) until this "controversey" hit the mainstream media, whereas there have been people refering to Archaeopteryx as a "feathered dinosaur" for decades. I use it every time I talk to the public on the subject, and no one has ever objected in the manner that school children pleaded for Pluto. Indeed, the idea that "birds ARE dinosaurs" has such strong appeal to the public that detracters used to accuse pro dino-bird scientists of pandering to the public (Larry Martin used to do this all the time). Restricting Aves (and "birds", frankly) only reinforces an idea that is already popular with the public while simultaneously opening a "teachable moment" to talk about evolution and how out groups get succesively more like the crown group the more closely related they are.

And finally, planets HAVE to be defined via platonic ideals, because they don't descend genetically from other plantes. You can hash out whether Pluto is similar enough in size, mineral composition, and formational history to other planets...that is, if it's "planety" enough, but it's emphatically NOT the same as classifying things by common descent.

Astronomers did it simply because it was more useful conceptually to their field. Not only is is useful to stop extending names in ways that lead to unjustified inferences about stem groups, but there is immense pedagological and conceptual value in teaching people to think this way. It would be tragic if we didn't have the constitutional fortitude to tread where our atronomer brethren have gone, especially withe the popular and educational advantages we have.

In short, I have yet to see any arguement in favor of defining Archaeopteryx into Aves a priori that doesn't boil down to "I'm used to it" (or perhaps, "a lot of people are used to it"), and while there is merit in such an argument, I don't think it supercedes the operational and educational benefits in this case.


Scott Hartman Science Director Wyoming Dinosaur Center 110 Carter Ranch Rd. Thermopolis, WY 82443 (800) 455-3466 ext. 230 Cell: (307) 921-8333

www.skeletaldrawing.com