Jon Barber wrote (in part):
D) It lacked an alula, which assists in low-speed flight and maneuverability control.<
It's been my hypothesis for a long time that the aerodynamic function of the alula could have been fulfilled by the exposed finger claws of early birds such as Archie (see my post from several years ago at http://dml.cmnh.org/2001Jan/msg00543.html).