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Re: Apatosaurus vs Brontosaurus
I would add that the taxonomy of the three _Apatosaurus_ species (_ajax_,
_excelsus_, louisae_) is currently unresolved.
In the early 1990's, McIntosh distinguished _ajax_ from _excelsus_ based on
overall size: _ajax_ has a larger adult body size than _excelsus_. However,
McIntosh also points out that, apart from "body size", the two species are
indistinguishable based upon the described material. Rather than splitting
_ajax_ and _excelsus_ into two separate genera and thus resurrecting the
genus _Brontosaurus_(which has _excelsus_ as its type species), it is more
likely that the two would be combined into one species.
The third species, _A. louisae_ (for which a skull is known), was previously
distinguished from the other two species based on vertebral characters
(forward expansion on the cervical ribs in _ajax_ and _excelsus_ and the
lack of this expansion in _ louisae_ ). However, the diagnostic nature of
this character has recently been called into question...
http://www.cmnh.org/dinoarch/2003May/msg00250.html
http://www.cmnh.org/dinoarch/2003Apr/msg00091.html
Wedel, M.J. and Sanders, R.K. (2002). Osteological correlates of cervical
musculature in Aves and Sauropoda (Dinosauria: Saurischia), with comments on
the cervical ribs of _Apatosaurus_. PaleoBios 22(3): 1-6.
Thus, it may be that all three _Apatosaurus_ species could be combined into
one species: _A. ajax_, with _excelsus_ and _louisae_ as junior synonyms.
At the moment, _A. yahnapin looks safe in its own genus (_Eobrontosaurus_).
As for _A. minimus_ (also from the Morrison Formation)... it as been mooted
as a possible titanosaur based on the presence of six sacrals and the shape
of the ilium. Unless previous descriptions of _A. minimus_ are seriously
awry, this species clearly does not belong in _Apatosaurus_.
Tim
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