[Date Prev][Date Next][Thread Prev][Thread Next][Date Index][Thread Index][Subject Index][Author Index]

Huaxiagnathus orientalis



Huaxiagnathus Hwang, Norell, Ji and Gao 2004
= "Huaxiasaurus" anonymous 2000
H. orientalis Hwang, Norell, Ji and Gao 2004
Middle Barremian, Early Cretaceous
Jianshangao Beds(?) of Yixian Formation, Liaoning, China
Holotype- (CAGS-IG02-301) (~1.6 m; subadult) nearly complete skeleton
missing caudal vertebrae past the 25th (radius 51 mm; metacarpal I 19 mm;
phalanx I-1 38 mm; metacarpal II 40 mm; phalanx II-1 26 mm; phalanx II-2 35
mm; metacarpal III 26 mm; femur 163 mm; metatarsal III 102 mm)
Referred- (NGMC 98-5-003; specimen of "Huaxiasaurus") (~1.8 m) partial
skeleton including skull, forelimb, hindlimb

I have the pdf, if anyone's interested.  I also have a photo of the referred
specimen that's not in the paper.  It's from Hwang et al.'s SVP poster, but
now that this is published, I don't see a problem with distributing it.
It's not like you can see anything in it anyway ;-)

The authors get the following topology-

222 characters, 55 taxa, strict consensus, 2592 mpt's
|--Allosaurus
`--+--Sinraptor
   `--+--+--Albertosaurus
      |  `--Tyrannosaurus
      `--+--+--Pelecanimimus
         |  `--+--Shenzhousaurus
         |     `--+--Harpymimus
         |        `--+--Garudimimus
         |           `--+--Archaeornithomimus
         |              |--Gallimimus
         |              |--Struthiomimus
         |              |--Ornithomimus
         |              `--Anserimimus
         `--+--Ornitholestes
            `--+--+--Huaxiagnathus
               |  `--+--Sinosauropteryx
               |     `--Compsognathus
               `--+--+--Alvarezsaurus
                  |  `--+--Patagonykus
                  |     `--+--Shuvuuia
                  |        `--Mononykus
                  |--+--+--Alxasaurus
                  |  |  `--+--Erlikosaurus
                  |  |     `--Segnosaurus
                  |  `--+--Incisivosaurus
                  |     `--+--Caudipteryx
                  |        `--+--+--Avimimus
                  |           |  |--Caenagnathidae
                  |           |  `--Microvenator
                  |           `--+--Oviraptor
                  |              |--Rinchenia
                  |              |--Citipati
                  |              |--Conchoraptor
                  |              `--Ingenia
                  `--+--+--Rahonavis
                     |  `--+--Archaeopteryx
                     |     `--Confuciusornis
                     `--+--+--Sinovenator
                        |  `--+--IGM 100/44
                        |     `--+--Byronosaurus
                        |        |--Sinornithoides
                        |        `--+--Troodon
                        |           `--+--Saurornithoides mongoliensis
                        |              `--Saurornithoides junior
                        `--+--Microraptor zhaoianus
                           |--Sinornithosaurus
                           `--+--Utahraptor
                              |--Saurornithoides
                              |--IGM 100/1015
                              |--Velociraptor
                              |--Deinonychus
                              |--Unenlagia
                              |--Adasaurus
                              |--Achillobator
                              `--Dromaeosaurus

I added Huaxiagnathus to my analysis, and used the supplementary info to
code several more taxa newly added to the TWG matrix with data not
previously in my analysis (Sinosauropteryx- 8 new codings; Compsognathus- 9;
Shenzhousaurus- 14; Archaeornithomimus- 16; Anserimimus- 10).  Particularly
interesting is that Anserimimus is coded for many pelvic and a couple
vertebral characters, these areas being otherwise completely undescribed in
the literature.

So, 243 characters, 112 taxa, majority rule consensus, 114901 mpt's
|--+--Allosaurus
|  `--+--Sinraptor
|     `--Monolophosaurus
`--+--Iliosuchus
   |--Stokesosaurus
   `--+--Fukuiraptor
      |--+--"Chilantaisaurus" maortuensis
      |  `--+--Eotyrannus
      |     `--Aviatyrannis
      `--+--+--Tyrannosauridae
         |  `--+--Dryptosaurus
         |     `--Bagaraatan
         `--+--Proceratosaurus
            `--+--Huaxiagnathus
               `--+--Calamosaurus
                  `--+--+--Sinosauropteryx
                     |  `--Nqwebasaurus
                     `--+--Compsognathus
                        `--+--Scipionyx
                           `--+--+--Coelurus
                              |  `--YPM 1996
                              `--+--Ornitholestes
                                 `--Maniraptoriformes

|--+--+--Therizinosaurus
|  |  `--+--+--Patagonykus
|  |     |  `--+--Alvarezsaurus
|  |     |     `--Mononykinae
|  |     `--+--+--Garudimimus
|  |        |  `--+--Archaeornithomimus
|  |        |     `--Ornithomimidae
|  |        `--+--Pelecanimimus
|  |           `--+--"Grusimimus"
|  |              `--+--+--Harpymimus
|  |                 |  `--Shenzhousaurus
|  |                 `--+--Deinocheirus
|  |                    `--Anserimimus
|  `--+--+--+--Beipiaosaurus
|     |  |  `--Nothronychus
|     |  `--+--Alxasaurus
|     |     `--+--Nanshiungosaurus
|     |        `--+--+--Erliansaurus
|     |           |  `--Neimongosaurus
|     |           `--+--Erlikosaurus
|     |              `--Segnosaurus
|     `--+--+--Caudipteryx zoui
|        |  `--+--Caudipteryx sp. nov.
|        `--+--+--Incisivosaurus
|           |  `--Avimimus
|           `--+--Nomingia
|              `--+--Caenagnathidae
|                 `--+--Oviraptoridae
|                    `--Microvenator
`--+--Yixianosaurus
   `--+--Protarchaeopteryx
      `--Deinonychosauria

|--+--Dromaeosaurus
|  `--+--Utahraptor
|     `--+--Pyroraptor
|        `--+--+--Bambiraptor
|           |  `--+--+--Sinornithosaurus
|           |     |  `--NGMC 91
|           |     `--+--Richardoestesia
|           |        `--+--Microraptor gui
|           |           `--+--Microraptor zhaoianus
|           |              `--Cryptovolans
|           `--+--IGM 100/1015
|              `--+--Saurornitholestes
|                 `--+--Adasaurus
|                    `--+--+--Unenlagia
|                       |  `--Deinonychus
|                       `--+--Achillobator
|                          `--Velociraptor
`--+--+--Archaeornithoides
   |  `--+--IGM 100/44
   |     `--+--Sinovenator
   |        `--+--Byronosaurus
   |           `--+--Sinornithoides
   |              `--+--Saurornithoides junior
   |                 `--+--Saurornithoides mongoliensis
   |                    `--Troodon
   `--+--Scansoriopterygidae
      `--+--+--Archaeopteryx
         |  `--Wellnhoferia
         `--+--Yandangornis
            `--+--+--Jixiangornis
               |  `--+--Rahonavis
               |     `--Shenzhouraptor
               `--+--+--Sapeornis
                  |  `--Omnivoropteryx
                  `--+--+--Confuciusornis
                     |  `--Changchengornis
                     `--+--Jibeinia
                        `--+--Longipteryx
                           `--+--Protopteryx
                              `--+--Concornis
                                 `--+--Catalan nestling
                                    `--+--Eocathayornis
                                       `--+--+--Sinornis
                                          |  `--+--Iberomesornis
                                          |     `--Longirostravis
                                          `--+--+--Eoalulavis
                                             |  `--Liaoningornis
                                             `--+--+--+--Boluochia
                                                |  |  `--Gobipteryx
                                                |  `--+--Noguerornis
                                                |     `--+--Enantiornis
                                                |        `--Neuquenornis
                                                `--+--+--Apsaravis
                                                   |  `--Ambiortus
                                                   `--+--+--Patagopteryx
                                                      |  `--Songlingornis
                                                      `--+--Yanornis
                                                         `--+--Yixianornis
                                                            `--Ornithurae

As an interesting experiment, I decided to run my analysis with only those
taxa used by Hwang et al.  Thus, the only taxon differences are they divide
up a few OTU's of mine (Tyrannosauridae, Ornithomimidae, Mononykinae,
Oviraptoridae), have Chirostenotes pergracilis instead of Caenagnathidae,
and combine the species of Caudipteryx into one OTU.

Strict consensus, 204 mpt's-
|--+--Sinraptor
|  `--Allosaurus
`--+--Tyrannosauridae
   `--+--Sinosauropteryx
      |--Huaxiagnathus
      |--Ornitholestes
      `--+--Compsognathus
         `--+--+--Anserimimus
            |  `--+--Pelecanimimus
            |     `--+--Harpymimus
            |        `--+--Shenzhousaurus
            |           `--+--Garudimimus
            |              `--+--Archaeornithomimus
            |                 `--Ornithomimidae
            `--+--+--+--Patagonykus
               |  |  `--+--Alvarezsaurus
               |  |     `--Mononykinae
               |  `--+--+--Alxasaurus
               |     |  `--+--Erlikosaurus
               |     |     `--Segnosaurus
               |     `--+--Avimimus
               |        `--+--+--Caudipteryx zoui
               |           |  `--Caudipteryx sp. nov.
               |           `--+--Incisivosaurus
               |              |--Caenagnathidae
               |              `--+--Microvenator
               |                 `--Oviraptoridae
               `--+--Sinovenator
                  |--IGM 100/44
                  |--+--Sinornithoides
                  |  `--+--Saurornithoides mongoliensis
                  |     `--+--Saurornithoides junior
                  |        `--Troodon
                  `--+--+--Microraptor
                     |  `--+--Byronosaurus
                     |     `--+--Confuciusornis
                     |        `--+--Rahonavis
                     |           `--Archaeopteryx
                     `--+--Utahraptor
                        `--+--Sinornithosaurus
                           `--+--Saurornitholestes
                              |--IGM 100/1015
                              |--Unenlagia
                              |--Velociraptor
                              |--Adasaurus
                              `--+--Deinonychus
                                 `--+--Dromaeosaurus
                                    `--Achillobator

Interesting how which taxa are included affects topology, isn't it?  Another
example of why it's important to include all the taxa you can.

Mickey Mortimer