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RE: Dromornithids and size limits.



From: "Jaime A. Headden" <qilongia@yahoo.com>
Some birds do precisely that, standing on one leg and using the other to
play with things.

Are these ground or arboreal birds? Being able to use one foot while the other is tightly gripping a tree branch isn't quite the same as _balancing_ on one leg while using the other.



However, parrots and turacos are birds that are effectively three-legged climbers, in that they use >their beaks to manipulate and clamb with.

Effectively, yes. But are they _effective_ climbers? Aren't most tree-adapted quadrupeds likely to be faster, and more agile? Plus, they have their jaws free. Can a parrot climb and hold a nut in its beak at the same time? Well, maybe it could, but it'd still be clumsy.


Many corvoids and parrots use tools held in the beak.

Quite true. But _how_much_better_ can they improve? Take, as representatives, a sparrow and a mouse. Subject them to intense selection pressure for tool manipulation. The sparrow will come to do a _lot_ with its beak, but I doubt it'll be able to match the mouse with its two limbs of manipulators.


Birds that need to manipulate develop a means, usually in the head.

They certainly do. All I'm saying is that a tetrapod is, all things being equal, better poised to develop a _superior_ means. Birds usual maintain their edge despite "inferior" adaptations precisely because of their wings. Lose flight and get stuck with the "inferior". Some niches _will_ remain open for obligate terrestrial birds, but they don't have the evolutionary _pathways_ to explore as many niches as tetrapods...or at least, not explore them as quickly and effectively as a tetrapod itself.


If they were to lose flight, for instance, and specialize as fossorial, it is likely that they...would
specialize for it. The need to specialize for burrowing is, to a point, unneccesary.

In their current niches, yes. If a flightless bird _attempted_ to enter the fossorial niche, all things being equal, it would find itself at a great disadvantage against a tetrapod entering that niche _at_the_same_time. Because a generalized tetrapod is already a better excavator than a generalized biped. If you're dealing with a flightless bird who's been so for some time, so that the wings have atrophied to almost nothing (like kiwis), it's at an even _bigger_ disadvantage because bringing the wings into play as shovels requires bringing them back as _limbs_ in the first place.


Meanwhile, larger birds like phorusrhacoids have HUGE heads for rending...

Meanwhile, while the Killer Bird has one predatory appendage, the sabretooth, or other predator has _three_. A lion leaping up to claw and bite as the hindquarters of its fleeing prey has a better chance of staying attached than a phorusrhacoid would with its single, albeit wickedly hooked, anchor into the prey's flesh. _In_general_ a tetrapod predator has an advantage over a biped predator.


No mammal niche is threatened by birds.

Exactly. I think we might actually agree here. It's just a matter of interpretation. You say they _aren't_ threatened. I say they _can't_ be threatened. Because, within those niches, tetrapods are better suited to develop the requisite adaptations.


I say nothing against the marvelous talents of birds. They rule the skies with an awesome diversity and fecundity. I merely say that, having specialized to such an amazing degree, they pay the price of finding it more difficult to de-specialize and find something else. They are, in a sense, stuck.

That's all.

Eric
______________________________________________________________________
"There is no other wisdom,
And no other hope for us
But that we grow wise. -- Diane Duane
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