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Re: New refs (long)
<Are you kidding???!!!>
I never write 3 page emails just to kid ;)
<I'll list just a few "extant animal groups" that display a range of
metabolisms:
Mammals: <snip> Sharks: <snip> Fishes: <snip> Insects: <Snip>>
Ok, this is a much better line of evidence than claiming that big mammals
(all of which are placentals today) are less endothermic than small mammals.
Itâs inconsistent, though, to use so many marine organisms as your examples
of metabolic diversity, but then argue, âno extant land mammal has ever
achieved the dimensions of the largest sauropods - or even the largest
hadrosaurs.â
<All your math pertains to placental mammals. You're assuming (very bravely)
that Placentalia constitute a satisfactory analog for the Dinosauria.
However, no extant land mammal has ever achieved the dimensions of the
largest sauropods - or even the largest hadrosaurs. Your choice of the
placental clade of mammals for comparison is a Strawman.>
Ok, let me get this straightâarthropods and lamniforms are good dinosaur
analogs, but placental mammals are a strawman?! I think your comparative morph
maybe a bit rusty. To begin, the math applied to all terrestrial
tetrapodsâthe constant of 70 kcal, in 70 kcal / kg^.75 applies as a general
rule to placental mammals. No extant land mammal gets close to the largest
sauropods, but extinct indricotheres get as large as many sauropods, and are
larger than any known hadrosaurs skeleton (I understand there is ichnological
data suggestive of larger hadrosaurs). Do you suggest that indricotheres were
ectothermic? For that matter, how about probiscideans, since most
non-sauropods donât get larger than big bull _Loxidonta_?
<Placental mammals represent a very limited (and very artificial) clade for
comparison to dinosaurs. Would you say that _Coelophysis_ and _Brachiosaurus_
had the same metabolism as any modern bird? If you don't, then you're argument
is already dead in the water. If you do, then you're on your own...>
Iâm really not sure what you mean by placental mammals being an
artificial clade. As in not monophyletic? Iâm sure I could find a few
mammals paleontologists who would take issue with that. Surely you arenât
referring to the amount of morphological diversity? It far exceeds that of
dinosaurs. There are no brachiating dinosaurs, no cetacean analogs, no sea
otter analogs. Even ignoring pinnipeds, Carnivora has far more diversity in
its lineage than theropods (exclusive of aves). There is far more variety in
niche exploitation and locomotive style in placentals than in dinosaurs.
Youâre right, of course, that inclusion of marsupials and monotremes
greatly expands the range of metabolic adaptation seen in Mammalia, but then
you are including groups with three different reproductive styles, for crying
out loud! I would be very interested indeed in arguments that claim that
dinosaurs are more diverse than the âartificialâ group of placental
mammals. Or why arthropods or chondrithyans are better models.
As for your final question, âWould you say that _Coelophysis_ and
_Brachiosaurus_ had the same metabolism as any modern bird?â Yes. A kiwi
has a RMR of 40 kcal / kg^.75. Much higher than the 5-15 kcal range of extant
reptiles, much lower than the 70 of mammals. Based on the growth style in
sauropods and early theropods, which exceeds that of âgigantothermicâ
Deinosuchus, it seems reasonable to suppose that both genera have RMRs at least
that high (especially in light of data that shows sauropod juveniles growing
faster than other dinosaursâdonât anyone say itâs because they have to
grow more to attain large size, thatâs not what Deinosuchus does!).
<The fact that some dinosaurs were feathered and (as far we can tell) all
dinosaurs were not is positive evidence for metabolic variation within the
Dinosauria.>
Like the fact that some placentals were furry and others (most assuredly) are
not proves that some placentals had a reptilian RMR? This line of reasoning
may prove telling; one can compensate with fat deposits and behavioral
regulation, but if we find a few unfeathered hypsys, etc, I would admit that it
is evidence in favor of your supposition, though not conclusive.
Scott