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"Cetiformes" and Dinosauriformes




Nick and Zoe,
Actually I should have put the name in quotation marks: "Cetiformes", since this name has never been proposed in the literature AFAIK.
Even though I am more of a lumper than a splitter (in general), in this case, I felt that there was a definite possibility that whales could be polyphyletic (diphyletic). Therefore in my 1994 book, I split them into three separate Orders: Archaeocetiformes (7 families), Odontocetiformes (17 families), and Mysticetiformes (6 families). I coded them as a clade, and if its holophyly can be clearly demonstrated, I would have no problem combining them into one Order "Cetiformes". I don't see any reason to use the spelling "Cetaciformes" (which I assume is based on a plural form "cetacea" rather than the singular "cetus"). If there is a linguistic problem here, I would like to hear it (but if it is just a phylocode thing, I am not going to give it much weight).
I coded Order Mesonychiformes as sister group to the whale clade, but recent evidence clearly shows that many (if not all) whales are closer to Order Artiodactyliformes. Therefore, as I said, I plan to move Mesonychiformes up to show them as sister group to an artiodactyl-whale clade (cetartiodactyls) and still assume that whales are probably holophyletic.
However, if whales do prove to be diphyletic, I will then move up any of the three whale Orders up next to Mesonychiformes if need be, and would of course abandon any contingency plans to formally recognize a "Cetiformes". I don't think the non-holophyly possibility has been thoroughly tested by morphologists or the molecularists.
This is exactly the same approach that I took with the dinosaurs, with Order Saurischiformes and Order Ornithischiformes coded as a "dinosaur" clade. If and when I am satisfied that there is a set of significant dinosaur "synapomorphies", I will combine them into a single Order Dinosauriformes, which will probably closely match the content of the cladistically defined Dinosauriformes. I prefer to "lump", but on the other hand I only formally recognize (with names) the most strongly supported clades.
I am still studying some of the more "significant" dinosaur synapomorphies that have been suggested: astragalar ascending process, cervical epipophyses, large deltopectoral crest, some form of partially open acetabulum, elongate vomers, and perhaps even the reduced posttemporal openings (although the latter has arisen independently in a lot of other groups). Thus the latter one may end up off of my list (along with "three or more sacral vertebrae", twist thumb, and postfrontal absent). The "three sacral vertebrae" thing was what really got me into dinosaur origins, and as far as I'm concerned, Galton has laid that one to rest. You can call that "synapomorphy" anything you want (bad, insignificant, weak, rejected, or whatever), it's definitely off my list, and I am concentrating on those that seem to be potentially better (more significant, stronger, more worthwhile, etc.). If that is "subjective" weighting, so be it. That's the approach that I personally find most valuable (see Mayr and Ashlock's 1991 book for a discussion of discrimating between characters and what types should be given more weight).
-------Ken Kinman
*******************************************
From: NJPharris@aol.com
To: <kinman@hotmail.com>, <dinosaur@usc.edu>
Subject: Re: Whales, not whippos (Probably!!?)
Date: Fri, 21 Sep 2001 14:57:40 EDT

In a message dated Fri, 21 Sep 2001 1:44:04 PM Eastern Daylight Time, "Ken Kinman" <kinman@hotmail.com> writes:

> This new information makes me even more determined to maintain a
> separate Order Mesonychiformes, and (for the time being) coded as sister
> group to a cetartiodactyl clade---- but with artiodactyls and whales still
> as separate orders from one another and the latter as exgroup of the former:
>
> 14 Mesonychiformes
> 15 Artiodactyliformes
> _a_ Cetiformes


Shouldn't this last one be "Cetaciformes"?

--Nick P.


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