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Re: dinosaur synapomorphies? (posttemporal opening)



Anybody can take two taxa and use them to define a holophyletic clade. The question is whether they should. Clades should be named for groups that appear to have strong synapomorphies, and to me "postfrontals absent" cannot be defended as strong. A long list of weak characters isn't convincing to me, especially when reversals are automatically invoked whenever the weak characters are not congruent.
Question. If the three dinosaur "synapomorphies" not found in sauropods aren't reversals, and they lie outside the Triceratops-bird defined "Dinosauria", do we then "educate" traditionalists and the public that sauropods are not dinosaurs, or do we redefine Dinosauria?
And I still haven't gotten any good answers to which of Sereno's dinosaur synapomorphies are the strongest (clearly separate dinosaurs from non-dinosaurs). Just two or three (is the long deltopectoral crest pretty reliable?) I've already criticized "postfrontals absent".
What about posttemporal opening reduced to small foramen? Compared to what? How small is small? Are there no "small" ones in any basal crurotarsans? Someone posted that this sounds like a strong synapomorphy, but admitted not even knowing what it is. Was that supposed to be convincing?? Is there anyone who knows more about it who wants to defend this synapomorphy? Aren't pterosaur and pelycosaur posttemporal openings small? I see no reason to trust such a character.
-----Ken Kinman
*******************************************
From: Dinogeorge@aol.com
Reply-To: Dinogeorge@aol.com
To: dinosaur@usc.edu
Subject: Re: dinosaur synapomorphies?
Date: Tue, 28 Aug 2001 00:42:17 EDT

In a message dated 8/27/01 11:44:25 AM EST, kinman@hotmail.com writes:

<< It amazes me that so many people now take the holophyly of Dinosauria for
granted. >>


Holophyly of Dinosauria is now legislated by its definition, which is
node-based: the common ancestor of modern birds and Triceratops and all its
descendants. This is beyond taking it for granted. I'm not too thrilled by
this definition and for historical reasons would have preferred this one: the
common ancestor of Megalosaurus and Iguanodon and all its descendants. This
is, as far as we know, the exact same clade (though it need not be, for
example, in the unlikely event that birds are not descended from this common
ancestor). According to the Triceratops-birds definition, all birds are
dinosaurs, period. It's not even an issue any more.


Definitions of clades are stated in terms of relationships, not in terms of
characters or features; today's synapomorphies are tomorrow's homoplasies,
and character-based definitions are likewise subject to instability. What
makes any animal a dinosaur is simply that it descended from the common
ancestor of birds and Triceratops. What might allow us to determine whether a
particular animal is a dinosaur are its anatomical characters and features,
which support or refute its hypothesized descent from that common ancestor.
If it shares enough features with animals that are already known to belong to
this elite clade (such as Triceratops and modern birds, which are dinosaurs
by definition), then it is likely that it is a dinosaur as well.


You should frame your critiques of dinosaur features keeping these ideas in
mind.


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