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Bipedalism in the trees
Bipedalism evolving on the ground and moving into the trees or bushes
seems reasonable, more so to me than bipedalism evolving in the trees in
the first place. I think the arguments about extant tree-dwellers and their
use of all four limbs are good ones.
Ground-originating bipedalism would provide the putative creature with
the coordination and balance necessary to be able to perch on a branch
as a biped. The forelimbs, which may have evolved for prey capture,
could have been modified to preserve prey capture but also provide
additional balance, in the same way that a tightrope walker uses arms or
a pole. In this scenario, lengthening the arms would be good for balance.
The lengthened arms, possibly covered already with feathers or
proto-feathers, could then have been used not just for balance but to
provide a controlled descent (pouncing on prey, jumping to another
branch), lift (for gliding from one perch to another), and eventually thrust.
JM Norton